Tortricidae
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  Pammene argyrana (Black-bordered Piercer)    Photo © Patrick Clement

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Tortricidae

3 Subfamilies.

Tortricinae
Wingspan 14-24mm. Medium sized broad winged moths. They hold the wings flat over their bodies, and have a bell shaped outline at rest, caused by a very convex base to the costa, which is then often rather distally excavated with a variably prominent apex producing a typical shape best seen in the genera Pandemis, Archips and Acleris. The forewing pattern consists of a darker basal patch, an outwardly oblique median fascia, and a subapical blotch. This typical pattern is however obscured in some species. Most of their larvae feed concealed in rolled or joined leaves or in spun shoots (from which they get the family name, Tortrix, meaning twister), and are often quite polyphagous on the foliage of various trees and shrubs, including conifers. Some species feed on herbaceous plants, some are more oligophagous and a few feed on fruits, berries, flowers and seed heads. Most overwinter as larvae and complete their feeding in the spring. Imagines of some of the genus Acleris hibernate and some are very polymorphic. Many fly actively in the late afternoon and evening or at dusk and sometimes come to light traps. They may often be disturbed in the day.

Chlidanotinae
Wingspan 11-16mm Only two species classified separately because of significant genitalia differences but otherwise with similar life histories to the Tortricinae.

Cochylini Somewhat smaller moths which have a characteristic resting posture with the closed wings forming a sharp ridge, and are often coloured white, yellow, pink or orange with variable darker markings, sometimes including metallic spots or striae. Their larvae feed internally in seed heads, roots, shoots and stems, mostly of herbaceous plants but some feed in berries and fruits of shrubs and small trees. Some are active in the day as well as later in the evening.

Olethreutinae
Wingspan 11-16mm. with a few as small as 9-11mm. or as big as 16-24mm. Small to medium sized moths with broad wings and a straighter costa than the Tortricini and whose wing markings and life histories are more specialised and varied. Some have similar wing patterns but typically they have a pale or dark dorsal blotch with white or pale costal marks (strigulae) and a circular marking at the tornus, often containing metallic scales, called an ocellus. There are however many variations on this basic scheme and some are polymorphic. Their larvae have very varied life histories, feeding in spinnings in leaves, shoots, flowers and seed heads and internally in stems, buds, roots, berries, seedpods, fruits and cones. They tend to be oligophagous or monophagous, and often feed on herbaceous plants. Some species are bivoltine. A few are diurnal and fly in sunshine but most are active in late afternoon and evening. Some come to light and some can be found resting on tree trunks in the day.

 Chlidanotinae
49.001 1013 Olindia schumacherana
White-barred Tortrix
49.002 1014 Isotrias rectifasciana
Hedge Tortrix
 Tortricinae
49.004 1010 Ditula angustiorana
Red-barred Tortrix
49.005 1006 Epagoge grotiana
Brown-barred Tortrix
49.008 1008 Philedone gerningana
Cinquefoil Tortrix
49.009 1007 Capua vulgana
Common Tortrix
49.010 1009 Philedonides lunana
Heath Twist
49.011 976 Archips oporana
Pine Tortrix
49.013 977 Archips podana
Large Fruit-tree Tortrix
49.014 979 Archips crataegana
Brown Oak Tortrix
49.015 980 Archips xylosteana
Variegated Golden Tortrix
49.016 981 Archips rosana
Rose Tortrix
49.017 982 Choristoneura diversana
Forest Tortrix
49.018 983 Choristoneura hebenstreitella
Great Tortrix
49.020 974 Argyrotaenia ljungiana
Heather Tortrix
49.021 987 Ptycholomoides aeriferana
Yellow Larch Tortrix
49.022 1000 Ptycholoma lecheana
Brindled Tortrix
49.023 971 Pandemis cinnamomeana
White-faced Tortrix
49.024 969 Pandemis corylana
Chequered Fruit-tree Tortrix
49.025 970 Pandemis cerasana
Barred Fruit-tree Tortrix
49.026 972 Pandemis heparana
Dark Fruit-tree Tortrix
49.028 986 Syndemis musculana
Dark-barred Tortrix
49.029 1002 Lozotaenia forsterana
Large Ivy Tortrix
49.030 985 Cacoecimorpha pronubana
Carnation Tortrix
49.031 989 Zelotherses paleana
Timothy Tortrix
49.032 990 Zelotherses unitana
Northern Grey Tortrix
49.033 988 Aphelia viburnana
Bilberry Tortrix
49.035 991 Clepsis senecionana
Obscure Tortrix
49.037 993 Clepsis spectrana
Cyclamen Tortrix
49.038 994 Clepsis consimilana
Privet Tortrix
49.039 998 Epiphyas postvittana
Light Brown Apple-moth
49.040 1001 Lozotaeniodes formosana
Orange Pine Tortrix
49.041 999 Adoxophyes orana
Summer Fruit Tortrix
49.042 1027 Neosphaleroptera nubilana
Deep Brown Shade
49.043 1026 Exapate congelatella
Autumnal Shade
49.044 1025 Tortricodes alternella
Winter Shade
49.045 1029 Eana osseana
Dotted Shade
49.047 1030 Eana incanana
Bluebell Shade
49.049 1024 Cnephasia incertana
Light Grey Tortrix
49.050 1020 Cnephasia stephensiana
Grey Tortrix
49.051 1021 Cnephasia asseclana
Flax Tortrix
49.052 1022 Cnephasia pasiuana
Meadow Shade
49.053 1022a Cnephasia pumicana
Cereal Tortrix
49.054 1023 Cnephasia genitalana
Dover Shade
49.055 1018 Cnephasia communana
May Shade
49.056 1019 Cnephasia conspersana
Coast Shade
49.057 1016 Cnephasia longana
Long-winged Shade
49.058 1034 Spatalistis bifasciana
Small Purple Button
49.059 1033 Tortrix viridana
Green Oak Tortrix
49.060 1032 Aleimma loeflingiana
Yellow Oak Button
49.061 1037 Acleris holmiana
White-triangle Button
49.062 1036 Acleris forsskaleana
Maple Button
49.063 1035 Acleris bergmanniana
Yellow Rose Button
49.064 1040 Acleris caledoniana
Caledonian Button
49.065 1039 Acleris comariana
Strawberry Tortrix
49.0655 1038x Acleris laterana/comariana
49.066 1038 Acleris laterana
Dark-triangle Button
49.067 1059 Acleris abietana
Perth Button
49.069 1041 Acleris sparsana
Ashy Button
49.070 1042 Acleris rhombana
Rhomboid Tortrix
49.071 1062 Acleris emargana
Notch-wing Button
49.073 1047 Acleris schalleriana
Viburnum Button
49.075 1052 Acleris umbrana
Dark-streaked Button
49.076 1054 Acleris cristana
Tufted Button
49.077 1048 Acleris variegana
Garden Rose Tortrix
49.078 1043 Acleris aspersana
Ginger Button
49.080 1053 Acleris hastiana
Sallow Button
49.082 1055 Acleris hyemana
Heath Button
49.083 1044 Acleris ferrugana
Rusty Oak Button
49.0835 1044x Acleris ferrugana/notana
49.084 1045 Acleris notana
Rusty Birch Button
49.085 1050 Acleris kochiella
Elm Button
49.086 1051 Acleris logiana
Grey Birch Button
49.087 1061 Acleris literana
Lichen Button
49.090 1015 Eulia ministrana
Brassy Tortrix
49.091 1011 Pseudargyrotoza conwagana
Yellow-spot Tortrix
49.092 921 Phtheochroa inopiana
Plain Conch
49.093 922 Phtheochroa schreibersiana
Scarce Gold Conch
49.094 923 Phtheochroa sodaliana
Buckthorn Conch
49.095 925 Phtheochroa rugosana
Rough-winged Conch
49.096 924 Hysterophora maculosana
Bluebell Conch
49.097 936 Cochylimorpha straminea
Straw Conch
49.099 933 Phalonidia gilvicomana
Wall-lettuce Conch
49.100 934 Phalonidia curvistrigana
Golden-rod Conch
49.101 926 Phalonidia manniana
Water-mint Conch
49.108 930 Gynnidomorpha alismana
Water-plantain Conch
49.109 937 Agapeta hamana
Common Yellow Conch
49.110 938 Agapeta zoegana
Knapweed Conch
49.111 954 Eupoecilia angustana
Marbled Conch
49.112 955 Eupoecilia ambiguella
Vine Moth
49.113 952 Commophila aeneana
Orange Conch
49.114 941 Aethes hartmanniana
Scabious Conch
49.115 942 Aethes piercei
Devil's-bit Conch
49.120 947 Aethes smeathmanniana
Yarrow Conch
49.121 939 Aethes tesserana
Downland Conch
49.122 949 Aethes dilucidana
Short-barred Yellow Conch
49.123 951 Aethes beatricella
Hemlock Yellow Conch
49.124 950 Aethes francillana
Long-barred Yellow Conch
49.127 945 Aethes cnicana
Thistle Conch
49.128 946 Aethes rubigana
Burdock Conch
49.129 959 Cochylidia rupicola
Hemp-agrimony Conch
49.130 958 Cochylidia subroseana
Dingy Roseate Conch
49.132 956 Cochylidia implicitana
Chamomile Conch
49.133 968 Thyraylia nana
Birch Conch
49.134 962 Cochylis roseana
Rosy Conch
49.135 963 Cochylis flaviciliana
Gold-fringed Conch
49.136 965 Neocochylis hybridella
White-bodied Conch
49.137 964 Neocochylis dubitana
Little Conch
49.138 964a Neocochylis molliculana
Ox-tongue Conch
49.139 966 Cochylichroa atricapitana
Black-headed Conch
49.142 960 Falseuncaria ruficiliana
Red-fringed Conch
 Olethreutinae
49.143 1114 Eudemis porphyrana
Apple Marble
49.144 1113 Eudemis profundana
Diamond-back Marble
49.145 1088 Pseudosciaphila branderiana
Great Marble
49.146 1089 Apotomis semifasciana
Short-barred Marble
49.148 1091 Apotomis lineana
Willow Marble
49.149 1092 Apotomis turbidana
White-shouldered Marble
49.150 1093 Apotomis betuletana
Birch Marble
49.151 1094 Apotomis capreana
Sallow Marble
49.152 1095 Apotomis sororculana
Narrow-winged Marble
49.153 1096 Apotomis sauciana
Bilberry Marble
49.154 1087 Orthotaenia undulana
Woodland Marble
49.155 1086 Hedya salicella
White-backed Marble
49.156 1083 Hedya nubiferana
Marbled Orchard Tortrix
49.157 1082 Hedya pruniana
Plum Tortrix
49.158 1084 Hedya ochroleucana
Buff-tipped Marble
49.159 1085 Hedya atropunctana
Shoulder-spot Marble
49.160 1065 Celypha rufana
Lakes Marble
49.161 1063 Celypha striana
Barred Marble
49.162 1064 Celypha rosaceana
Roseate Marble
49.165 1066 Celypha woodiana
Mistletoe Marble
49.166 1076 Celypha lacunana
Common Marble
49.167 1068 Celypha rivulana
Silver-striped Marble
49.172 1073 Phiaris schulziana
Large Marble
49.173 1075 Phiaris micana
Olive Marble
49.178 1070 Stictea mygindiana
Cowberry Marble
49.179 1080 Olethreutes arcuella
Arched Marble
49.180 1079 Piniphila bifasciana
Pine Marble
49.181 1105 Lobesia occidentis
Spurge Marble
49.183 1108 Lobesia abscisana
Smoky-barred Marble
49.184 1106 Lobesia reliquana
Oak Marble
49.185 1109 Lobesia littoralis
Shore Marble
49.186 1097 Endothenia gentianaeana
Teasel Marble
49.187 1098 Endothenia oblongana
Downland Marble
49.188 1099 Endothenia marginana
Bordered Marble
49.189 1100 Endothenia pullana
Woundwort Marble
49.190 1101 Endothenia ustulana
Bugle Marble
49.191 1102 Endothenia nigricostana
Black-edged Marble
49.192 1103 Endothenia ericetana
Heath Marble
49.193 1104 Endothenia quadrimaculana
Blotched Marble
49.194 1111 Bactra lancealana
Rush Marble
49.195 1110 Bactra furfurana
Mottled Marble
49.199 1217 Eucosmomorpha albersana
Honeysuckle Bell
49.200 1216 Enarmonia formosana
Cherry-bark Moth
49.201 1117 Ancylis unguicella
Broken-barred Roller
49.202 1118 Ancylis uncella
Bridge Roller
49.203 1123 Ancylis laetana
Aspen Roller
49.204 1122 Ancylis obtusana
Small Buckthorn Roller
49.205 1116 Ancylis comptana
Little Roller
49.206 1121 Ancylis upupana
Dark Roller
49.207 1119 Ancylis geminana
Festooned Roller
49.209 1119a Ancylis diminutana
Small Festooned Roller
49.210 1125 Ancylis unculana
Buckthorn Roller
49.211 1128 Ancylis myrtillana
Bilberry Roller
49.212 1129 Ancylis apicella
Hook-tipped Roller
49.214 1126 Ancylis badiana
Common Roller
49.215 1115 Ancylis achatana
Triangle-marked Roller
49.216 1120 Ancylis mitterbacheriana
Red Roller
49.217 1124 Ancylis tineana
Rannoch Roller
49.220 1162 Rhopobota myrtillana
Bilberry Bell
49.222 1158 Rhopobota ustomaculana
Northern Bell
49.223 1159 Rhopobota naevana
Holly Tortrix
49.224 1205 Spilonota ocellana
Bud Moth
49.225 1205a Spilonota laricana
Larch-bud Moth
49.228 1153 Epinotia sordidana
Brown Alder Bell
49.229 1154 Epinotia caprana
Large Sallow Bell
49.230 1151 Epinotia trigonella
White-blotch Bell
49.231 1155 Epinotia brunnichana
Large Birch Bell
49.232 1152 Epinotia maculana
Dark Aspen Bell
49.233 1156 Epinotia solandriana
Variable Bell
49.234 1150 Epinotia abbreviana
Brown Elm Bell
49.237 1144 Epinotia signatana
Black-brindled Bell
49.238 1147 Epinotia cruciana
Willow Tortrix
49.239 1148 Epinotia mercuriana
Moorland Bell
49.240 1136 Epinotia immundana
Common Birch Bell
49.242 1145 Epinotia nanana
Small Spruce Bell
49.243 1135 Epinotia demarniana
Birch Bell
49.244 1132 Epinotia subocellana
White Sallow Bell
49.245 1137 Epinotia tetraquetrana
Square-barred Bell
49.246 1130 Epinotia pygmaeana
Dingy Spruce Bell
49.247 1131 Epinotia subsequana
Dark Spruce Bell
49.248 1139 Epinotia tenerana
Nut-bud Moth
49.249 1134 Epinotia ramella
Small Birch Bell
49.250 1140 Epinotia nigricana
Fir Bell
49.251 1146 Epinotia rubiginosana
Pine Bell
49.252 1142 Epinotia tedella
Common Spruce Bell
49.253 1143 Epinotia fraternana
Silver-barred Bell
49.254 1133 Epinotia bilunana
Crescent Bell
49.255 1138 Epinotia nisella
Grey Poplar Bell
49.256 1138a Epinotia cinereana
Grey Aspen Bell
49.257 1166 Zeiraphera griseana
Larch Tortrix
49.259 1163 Zeiraphera ratzeburgiana
Spruce-bud Moth
49.260 1165 Zeiraphera isertana
Cock's-head Bell
49.261 1157 Crocidosema plebejana
Southern Bell
49.264 1202 Eucosma obumbratana
Two-coloured Bell
49.265 1201 Eucosma cana
Hoary Bell
49.266 1200 Eucosma hohenwartiana
Bright Bell
49.267 1200b Eucosma fulvana
Fulvous Bell
49.269 1197 Eucosma campoliliana
Marbled Bell
49.270 1194 Eucosma aemulana
Obscure Bell
49.275 1192 Eucosma conterminana
Pale Lettuce Bell
49.276 1190 Eucosma aspidiscana
Golden-rod Belle
49.277 1199 Eucosma pupillana
Eyed Bell
49.278 1171 Gypsonoma minutana
Brindled Shoot
49.279 1169 Gypsonoma dealbana
Common Cloaked Shoot
49.280 1170 Gypsonoma oppressana
Poplar Shoot
49.281 1168 Gypsonoma sociana
White Cloaked Shoot
49.283 1167 Gypsonoma aceriana
Rosy Cloaked Shoot
49.284 1186 Epiblema sticticana
Colt's-foot Bell
49.285 1184 Epiblema scutulana
Thistle Bell
49.286 1184a Epiblema cirsiana
Knapweed Bell
49.288 1183 Epiblema foenella
White-foot Bell
49.289 1187 Epiblema costipunctana
Ragwort Bell
49.290 1182 Epiblema turbidana
Butterbur Bell
49.292 1174 Notocelia cynosbatella
Yellow-faced Bell
49.293 1180 Notocelia tetragonana
Square-spot Bell
49.294 1175 Notocelia uddmanniana
Bramble Shoot
49.295 1178 Notocelia roborana
Summer Rose Bell
49.296 1179 Notocelia incarnatana
Chalk Rose Bell
49.297 1177 Notocelia rosaecolana
Common Rose Bell
49.298 1176 Notocelia trimaculana
Triple-blotched Bell
49.299 1208 Pseudococcyx posticana
Dark Pine Shoot
49.303 1206 Clavigesta sylvestrana
Small Pine Shoot
49.304 1207 Clavigesta purdeyi
Pine Leaf-mining Moth
49.305 1210 Rhyacionia buoliana
Pine Shoot
49.306 1211 Rhyacionia pinicolana
Orange-spotted Shoot
49.307 1212 Rhyacionia pinivorana
Spotted Shoot
49.309 1285 Dichrorampha plumbana
Lead-coloured Drill
49.310 1286 Dichrorampha sedatana
Dingy Drill
49.311 1287 Dichrorampha aeratana
Obscure Drill
49.312 1280 Dichrorampha consortana
Downland Drill
49.313 1279 Dichrorampha acuminatana
Sharp-winged Drill
49.314 1282 Dichrorampha sylvicolana
Sneezewort Drill
49.315 1281 Dichrorampha simpliciana
Round-winged Drill
49.316 1278 Dichrorampha sequana
Square-spot Drill
49.318 1284 Dichrorampha vancouverana
Gold-fringed Drill
49.319 1275 Dichrorampha flavidorsana
Narrow-blotch Drill
49.320 1274 Dichrorampha alpinana
Broad-blotch Drill
49.321 1273 Dichrorampha petiverella
Common Drill
49.322 1276 Dichrorampha plumbagana
Silver-lined Drill
49.323 1283 Dichrorampha montanana
Spike Marked Drill
49.324 1257 Cydia nigricana
Pea Moth
49.325 1255 Cydia ulicetana
Grey Gorse Piercer
49.329 1266a Cydia illutana
Larch Piercer
49.330 1269 Cydia conicolana
Pine-cone Piercer
49.332 1268 Cydia coniferana
Pine-bark Piercer
49.334 1267 Cydia cosmophorana
Scarce Pine Piercer
49.335 1254 Cydia strobilella
Spruce-seed Moth
49.336 1266 Cydia pactolana
Scarce Spruce Piercer
49.338 1261 Cydia pomonella
Codling Moth
49.339 1256 Cydia servillana
Sallow-shoot Piercer
49.341 1260 Cydia splendana
Marbled Piercer
49.342 1259 Cydia fagiglandana
Large Beech Piercer
49.3421 1269a Cydia injectiva
American Cone-piercer
49.343 1262 Cydia amplana
Vagrant Piercer
49.3431 1215 Thaumatotibia leucotreta
False Codling Moth
49.345 1219 Lathronympha strigana
Red Piercer
49.347 1241 Grapholita compositella
Triple-stripe Piercer
49.348 1243 Grapholita pallifrontana
Liquorice Piercer
49.349 1242 Grapholita internana
Dark Gorse Piercer
49.351 1252 Grapholita lunulana
Northern Crescent Piercer
49.354 1251 Grapholita jungiella
Vetch Piercer
49.3541 1248 Grapholita molesta
Oriental Fruit Moth
49.356 1249 Grapholita lobarzewskii
Kent Fruit Piercer
49.357 1247 Grapholita funebrana
Plum Fruit Moth
49.358 1246 Grapholita tenebrosana
Deep-brown Piercer
49.359 1245 Grapholita janthinana
Pale-bordered Piercer
49.360 1223 Pammene splendidulana
Drab Oak Piercer
49.361 1228a Pammene ignorata
Devon Piercer
49.362 1227 Pammene giganteana
Early Oak Piercer
49.363 1228 Pammene argyrana
Black-bordered Piercer
49.364 1230 Pammene suspectana
Ash-bark Piercer
49.365 1229 Pammene albuginana
Blotched Piercer
49.366 1225 Pammene obscurana
Obscure Birch Piercer
49.367 1236 Pammene fasciana
Acorn Piercer
49.371 1239 Pammene rhediella
Fruitlet-mining Tortrix
49.372 1232 Pammene populana
Pigmy Y Piercer
49.373 1231 Pammene spiniana
Triangle-marked Piercer
49.374 1235 Pammene trauniana
Maple Piercer
49.375 1234 Pammene regiana
Regal Piercer
49.376 1233 Pammene aurita
Sycamore Piercer
49.377 1237 Pammene germmana
Black Piercer
49.378 1238 Pammene ochsenheimeriana
Black-patch Piercer
49.379 1272 Pammene aurana
Orange-spot Piercer
49.380 1271 Pammene gallicana
Purple-shaded Piercer
49.381 1221 Strophedra weirana
Little Beech Piercer
49.382 1222 Strophedra nitidana
Little Oak Piercer
   

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